003550000a220008500045001000016000002450152000165000045001680410013002137730043002261 aCook, David10aSexual selection in dung beetles I. A. multivariate study of the morphological variation in two species of Onthophagus (Scarabaeidae: Onthophagini) a0 aInglés1 tAust. Journal Zool.h32q123-132d1987